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We analysed 18S r RNA gene signatures in a sample collected below the chemocline of the anoxic Mariager Fjord in Denmark, and from these data we computed novel parametric and standard nonparametric estimates of protistan phylotype richness.

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Because anoxic environments have occurred throughout the Earth's history, the possibility remains that some little-known or even entirely unknown primitively anaerobic eukaryote groups still exist, and that anaerobic environments still hold early eukaryotes important for understanding the origin of the eukaryotic cell (Cavalier-Smith, 2004).

Until now this hypothesis has gone untested, because modern biodiversity research has barely touched the Earth's largest reservoirs of permanently anoxic systems (Dawson & Pace, 2002).

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